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The giant chameleon is a large lizard from the 1959 movie of Journey to the Center of the Earth. While it appears to be asleep while camouflaging, the lizard woke up and attacked the group, while they tried to escape by breaking a giant rock blocking their way using gunpowder left by Saknussem. After having its tongue stabbed, the lizard tried to attack the group (who were on a large sacrifical altar bowl), while it was clinging on a pillar, but it got knocked off of the pillar by a large rock and died by the running lava.
The giant chameleon does not look like an actual chameleon. Instead, it was portrayed by a painted Tegu lizard.